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*Leishmania* infection affects the immune system and the outcome of the disease may be determined by a fine balance of the regulatory pathways in a variety of ways. Studies analyzing gene expression in macrophages after *Leishmania* infection have revealed that induction of specific cytokines and changes in their receptor expression were the consequence of molecular pattern recognition^[@CR3],[@CR4],[@CR9],[@CR14]^. However, anti-inflammatory cytokines induced by *Leishmania* can contribute to disease progression^[@CR17]^. Therefore, balancing the pro- and anti-inflammatory responses is necessary for the successful resolution of the infection^[@CR22],[@CR23]^.

CCL2 is a mediator of macrophage migration and is important for the clearance of intracellular pathogens^[@CR15]^. In our study, CCL2 was induced upon infection with either *L. major* or *L. donovani* promastigotes or protozoan-antigen-pulsed dendritic cells. These data are consistent with those published previously^[@CR13]^. Activation of CCR2 with CCL2 was sufficient to confer the protective profile to macrophages upon *Leishmania* infection. Induction of CCR2 with CCL2 on *Leishmania*-infected macrophages resulted in the secretion of IL-12 and IL-10, which are cytokines required to control the *Leishmania*-infected macrophages^[@CR13]^. These results demonstrate that CCR2 is a critical element in the signaling cascade during *Leishmania* infection, inducing the protective cytokine IL-12 and decreasing IL-10 secretion, ultimately leading to the immune-mediated killing of intracellular pathogens. These data are consistent with our previous findings that CCR2 is required for macrophage activation
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